Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark white, cream, gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; symmetric at the base; not dissected; with small to large teeth on margin (teeth small). Epidermis not waxy. Venation pinnate. Secondary veins open, or closed; prominent. Smallest veins prominent, or not prominent, but visible; ladder like (veins close). Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant, or with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 3-6 (3-4 male, 3-6 female); free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 100 (many); filaments present; joined (base), or free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 3. Locules 3. Styles solitary. Stigmas 6 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 3 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves distinctly obovate; tertiary veins ladder-like and very close together; margins minutely toothed; inflorescences long racemose; styles long, and each one distinctly cleft.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.