Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark green, brown, black; not peeling in strips. Slash yellow, green, gray. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; ladder like. Intramarginal vein present to absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present, or represented by solitary flowers only; terminal, or axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across to to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present; the stamens inside the disk. Perianth present; with all whorls looking like tepals. Tepals 6. Stamens 3; filaments present; joined; free of the perianth; all equal in size; isomerous. Anthers 4 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles absent. Stigmas 9-12 lobed; distinctive; disk-like. Fruit capsular; fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); barely visible; wingless; uniform in cross-section; aril absent; endospermic.
Tepals persistent and recurved in fruit.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.