For my publications listed below, if you see the Acrobat PDF icon ( ), you may download the paper by right-clicking on the icon. If you instead see the grey icon ( ), the paper is not currently available for download. If you do not have Acrobat, you can download it free from Adobe.
Publications:
	Carlson, SC, DG Howarth, MJ Donoghue. (2011) Diversification of CYCLOIDEA-like genes in Dipsacacea (Dipsacales): implications for the evolution of capitulum inflorescences. BMC Evolutionary Biology 11(1): 325-337. [download PDF]
	Howarth DG, T Martins, E Chimney, MJ Donoghue. (2011) Diversification of CYCLOIDEA expression in the evolution of bilateral flower symmetry in Caprifoliaceae and Lonicera (Dipsacales). Annals of Botany 107(9): 1521-1532. [download PDF]
	Howarth DG, MJ Donoghue. (2009) Duplications and Expression of DIVARICATA-Like Genes in Dipsacales. Molecular Biology and Evolution 26(6): 1245-1258. [download PDF]
	Howarth DG, MJ Donoghue. (2006) Phylogenetic analysis of the “ECE” (CYC/TB1) clade reveals duplications predating the core eudicots. Proceedings of the National Academy of Sciences of the United States of America 103(24): 9101-9106. [download PDF]
	Howarth, DG, DA Baum. (2005) Genealogical evidence of homoploid hybrid speciation in an adaptive radiation of Scaevola (Goodeniaceae) in the Hawaiian Islands. Evolution 59(5): 948-961. [download PDF]
	Howarth, DG, MJ Donoghue. (2005) Duplications in CYC-like genes from dipsacales correlate with floral form. International Journal of Plant Sciences 166(3): 357-370. [download PDF]
	Randell, RA, DG Howarth, CW Morden. (2004) Genetic analyis of natural hybrids between endemic and alien Rubus (Rosaceae) species in Hawai’i. Conservation Genetics 5(2): 217-230. [download PDF]
	Howarth, DG, MHG Gustafsson, DA Baum, TJ Motley, (2003) Phylogenetics of the genus Scaevola (Goodeniaceae): Implication for dispersal patterns across the Pacific Basin and colonization of the Hawaiian Islands. (2003) American Journal of Botany 90(6): 915-923. [download PDF]
	Howarth, DG, DA Baum. (2002) Phylogenetic utility of a nuclear intron from nitrate reductase for the study of closely related plant species. Molecular Phylogenetics and Evolution 23(3): 525-528. [download PDF]
	Howarth, DG, DE Gardner, CW Morden. (1997) Phylogeny of Rubus subgenus Idaeobatus (Rosaceae) and its implications toward colonization of the Hawaiian Islands. Systematic Botany 22(3): 433-441. [download PDF]
Abstracts:
Howarth, D. G. and M. J. Donoghue. Phylogeny and expression of TCP and MYB genes and their role in floral symmetry shifts in core eudicots and Dipsacales. Botany Meetings 2008. Howarth, D. G. The Attraction of Flowers: Stories of their Evolution, Genes and Development. St John’s University Seminar, Feb. 2008. Howarth, D. G. and M. J. Donoghue. Phylogeny and expression of TCP and MYB genes and their role in floral symmetry shifts in Dipsacales, Botany Meetings 2007. Howarth, D. G. and M. J. Donoghue. Phylogeny and expression of TCP and MYB genes and their role in floral symmetry shifts in Dipsacales. New York Area Plant Molecular Biology Meeting, Yale University, June 2007. Howarth, D. G. and M. J. Donoghue. Evolution of floral symmetry genes (including CYCLOIDEA): Gene duplications and diversifying expression. Evolution Meetings 2006. Howarth, D. G. and M. J. Donoghue. Sequences from Dipsacales suggest deep eudicot duplications in the floral symmetry TCP genes. Botany Meetings 2004 Howarth, D. G. and M. J. Donoghue. Sequences from Dipsacales suggest deep eudicot duplications in the floral symmetry TCP genes. Evolution Meetings 2004. Howarth, D. G. and D. A. Baum. Genealogical evidence of homoploid hybrid speciation in an adaptive radiation of Scaevola (Goodeniaceae) in the Hawaiian Islands. Evolution Meetings 2004. Howarth, D. G. Evolution of Scaevola (Goodeniaceae), Bernice P. Bishop Museum special seminar, Hawaii. 2003. Howarth, D. G., and D. A. Baum. Comparing three variable nuclear intron regions and nrDNA ITS to determine the phylogenetic history and a possible hybrid origin in a closely related Hawaiian clade in Scaevola (Goodeniaceae). Botany 2002 (Abstract). Howarth, D. G. 2002. Evolution on Islands: Dispersal, Speciation, and Hybridization in Scaevola (Goodeniaceae) in the Hawaiian Archipelago. Department of Ecology and Evolutionary Biology special seminar, Yale University. Howarth, D. G., T. J. Motley, and M. H. G. Gustafsson. 1999. Phylogenetics and Biogeography of Scaevola in the Pacific Basin. International Botanical Congress XVI (Abstract).

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