Large trees to small trees. Buttresses present; tall, or short. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Slash brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; star-like. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation palmate. Secondary veins open; prominent. Smallest veins not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; terminal, or axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across to to about 3 cm long or across; round; with male or female parts raised on a column; with male and female parts raised on a stalk. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 8; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined, or partially joined, by base; 3-4. Locules 3-4. Styles free; 3-4. Stigmas 1 lobed; not distinctive. Fruit follicular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); much more than 1 cm long; winged; uniform in cross-section; aril absent; endospermic.
Leaves heart-shaped and with occasional stellate hairs; petals absent; fruits winged.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.