Large trees to small trees. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark red, gray, brown; not peeling in strips. Slash yellow, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; compound; pinnate; without a terminal leaflet; leaflets opposite, or leaflets alternate (rare); all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia present, or absent; pit-like, or appearing as hair tufts; scattered along midrib. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; terminal, or axillary, or on the trunk or branches; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant, or with male and female flowers on different plants; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-6; free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present, or absent; 1-5; free; not fleshy; deciduous; stalked; entire. Stamens 4-10; filaments present; free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 1-4. Locules 1-4. Styles solitary. Stigmas 2-4 lobed; not distinctive. Fruit capsular; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; less than 5 mm, or about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Leaflets generally opposite; petals well developed, usually with a bilobed scale, some (or all) may be reduced; usually only one carpel only developing; sarcotesta present; fruit covered in thick “hairs”. Taxon images: gennephe.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.