Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite, or whorled, or spiral; 4 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous to leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible, or not visible; net like, or ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units present, or absent; specialized inflorescence units umbels; numerous and persistent inflorescence bracts present at the bottom of the inflorescence. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 4. Stamens (4-)6(-8); filaments present; free of each other; free of the perianth; all equal in size; number of stamens not a multiple of the adjacent perianth whorl. Anthers 4 loculed; not awned; dehiscing by valves. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; distinctive; disk-like. Fruit berry-like; fleshy; not multiple; not splitting; enclosed in, or sitting on, the fleshy perianth, or not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Twigs with distinct scars from scales covering the leaf bud; bract involucre present; dioecious; anthers 4 celled. Taxon images: neoli.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Illustrations’ URL
http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz,
Paine and Zurcher (1993) should also be cited.