Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark green; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; star-like. Stinging hairs absent. Mature twig indumentum present; dense, or sparse.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; gland-dotted, or not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; terminal (rare), or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 4. Stamens 4; filaments present; free of each other; free of the perianth; all equal in size; isomerous. Anthers 2 loculed; not awned; dehiscing by long slits. Ovary superior. Carpels separate; 100 (many). Locules 1. Styles free. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves opposite; intramarginal vein present; oil glands absent; fruits on slender stalks. Taxon images: genmatt.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.