Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4; free; not fleshy; deciduous; not stalked; entire. Stamens 8; filaments present; free of each other; free of the perianth; of different sizes (alternating tall and short); diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 4. Locules 4. Styles solitary. Stigmas 4 lobed; distinctive; head shaped. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 4-8 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Young branchlets red-brown hairy; petioles swollen at the apex; flowers unisexual and 4-merous; fruit a 4-loculed drupe. Taxon images: maclur.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.