Emergent trees to shrubs, or climbers. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark red, gray, brown; not peeling in strips. Slash yellow, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds enclosed by leaves, or not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present, or absent. Stalk of blade absent to less than a quarter the length of the blade; winged, or not winged; attached to base of leaf blade; not swollen. Stipules present; joined; joined across twigs; encircling the twig; leafy; not fringed; large, or small; persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present, or represented by solitary flowers only; terminal; the flowers from a single point on the stem, or on an unbranched axis, or on a branched axis; specialized inflorescence units present, or absent; specialized inflorescence units heads; numerous and persistent inflorescence bracts absent. Flowers bisexual; not stalked, or stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; persistent in fruit, or not persistent in fruit; not growing any further after flowering, or growing slightly after flowering. Epicalyx absent. Petals present; 5; all joined; fleshy; deciduous; not stalked; entire. Stamens 5; filaments present; free of each other; joined to the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 1-2. Styles solitary. Stigmas 2 lobed; distinctive; head shaped. Fruit capsular, or berry-like; fleshy; not multiple; splitting, or not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); barely visible, or less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves opposite with interpetiolar stipules which from an ochrea early on, later splitting; secondary nerves barely visible. Taxon images: genfag2.gif genfagra.gif genfagel.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.