Large trees to small trees. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark white, gray; not peeling in strips. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent to not prominent, but visible. Smallest veins not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at both ends. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 5; filaments present; free of each other; joined to the perianth (sepals); all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2(-3). Locules 1. Styles free; 2(-3). Stigmas 1 lobed; distinctive; disk-like. Fruit capsular; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; winged. Seeds 1-2(-3) per fruit; about 1 cm long; wingless; uniform in cross-section; aril present; endospermic.
Stipules absent; fruit indehiscent; seeds with an scarlet aril. Taxon images: eleuth.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.