Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end (top). Stipules present, or absent; free from one another; laterally placed; not encircling the twig; scale-like; fringed, or not fringed; small; not persistent. Domatia present, or absent; pit-like; scattered along midrib. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary, or superaxillary; on an unbranched axis, or on a branched axis (E. brevipes); specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals (3-)4-5(-6); free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; (3-)4-5(-6); free; fleshy, or not fleshy; deciduous; stalked, or not stalked; lobed, or fringed like a comb. Stamens 6-60(-100) (many); filaments present; free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous, or triplostemonous, or polystemonous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; awned, or not awned; dehiscing by pores. Ovary superior. Carpels joined; 2-5. Locules 2-5. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy, or non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2-5 per fruit; less than 5 mm, or about 1 cm long; wingless; with folds like marble, in cross- section; aril absent; endospermic.
Leaves long petioled and kneed at the base of the lamina; petals toothed or fringed; anthers dehiscing by pores. Taxon images: elaeo.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.