Large trees. Buttresses absent. Spines absent. Stilt roots present. Bark gray, brown; not peeling in strips. Slash gray, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present (sometimes faint). Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked (short); about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals (4-)5; free, or partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; (4-)5; free; not fleshy; deciduous; stalked; entire. Stamens (4-)5; filaments present; free of each other; free of the perianth; all equal in size; isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary partly inferior. Carpels joined; (3-)4-5. Locules (3-)4-5. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 3-15 per fruit; less than 5 mm; winged; uniform in cross-section; aril absent; non-endospermic.
Large peat swamp forest tree with pneumatophores; petals distinctly stalked; seeds winged. Taxon images: dactyl.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.