Large trees to small trees. Buttresses present; short. Spines absent. Stilt roots absent. Bark brown; not peeling in strips. Slash orange, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy. Venation pinnate (rare), or trinerved. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 6 (two unequal whorls). Stamens 9 (third whorl has glands); filaments present; free of each other; free of the perianth; all equal in size; number of stamens not a multiple of the adjacent perianth whorl. Anthers (2-)4 loculed (2 in one species only, 4 in the rest); not awned; dehiscing by valves. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; much more than 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Large trees with buttresses; leaves opposite, glaucous, trinerved, with obvious tertiary veins; flowers small; fruits large. Taxon images: caryod.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.