Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark red, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate, or pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible, or not visible; ladder like (more like strands of hairs running across lamina). Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent, or persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; the flowers from a single point on the stem, or on an unbranched axis; specialized inflorescence units present, or absent; specialized inflorescence units umbels; numerous and persistent inflorescence bracts present at the bottom of the inflorescence, or absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; persistent in fruit; growing slightly after flowering. Epicalyx absent. Petals present; 5(-10); free; not fleshy; deciduous; not stalked; entire. Stamens 10(-100) (many); filaments present; free of each other; free of the perianth; all equal in size; isomerous, or polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary superior. Carpels partially joined, by styles; 5(-10). Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
No intramarginal vein; numerous bracts at the base of the condensed inflorescence; stipules not joined. Taxon images: bracken.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.