Emergent trees to small trees. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash red. Vertical stripes in the inner bark or wood absent. Odor absent, resinous, aromatic, garlic-like. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; compound; pinnate; without a terminal leaflet, or with a terminal leaflet; leaflets opposite, or leaflets alternate; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia present, or absent; pit-like; scattered along midrib. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts raised on a column, or not raised on a column; with male and female parts raised on a stalk. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5(-6); free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5(-6); free; not fleshy; deciduous; not stalked; entire. Stamens 5(-6); filaments present; free of each other; free of the perianth; all equal in size; isomerous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 5. Locules 5. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); about 1 cm long; winged; uniform in cross-section; aril absent; endospermic.
Large monoecious trees; bark fissured or shaggy; slash generally aromatic; leaves compound; seeds winged. Taxon images: toona.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.