Small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark red, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite, or spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation trinerved. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 6. Stamens 9; filaments present; free of each other; free of the perianth; all equal in size; number of stamens not a multiple of the adjacent perianth whorl. Anthers 4 loculed; not awned; dehiscing by valves. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; enclosed in, or sitting on, the fleshy receptacle; wingless. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Leaves trinerved; fruits with persistent tepals; anthers 4-loculed.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.