Large trees to shrubs. Buttresses present, or absent; short. Spines absent. Stilt roots present, or absent. Bark red, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; leathery, or fleshy; not gland-dotted; symmetric at the base; not dissected; smooth margined to with small to large teeth on margin (sometimes minutely gland tipped at margin). Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible, or not visible. Smallest veins not visible. Intramarginal vein absent. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; terminal, or axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on the same plant; not stalked, or stalked; about 0.5 cm long or across to to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present (at best very much reduced), or absent; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5(-8); free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens (5-)10; filaments present; free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous (usual). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 2-5 (reported for family). Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit drupaceous; fleshy to non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Mangrove trees, either buttresses or with pneumatophores; leaves spiral and subsessile; calyx tube with large bracteoles attached. Taxon images: lumnitz.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.