Small trees to shrubs, or climbers. Buttresses absent. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation trinerved to palmate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade to more than half the length of the blade; not winged; attached to surface of leaf blade, attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 3-6 (3-5(-6) in males, 4-6 in females). Stamens 3-5(-6); filaments absent; free of the perianth; all equal in size; isomerous. Anthers 2 loculed; not awned; dehiscing by valves. Ovary inferior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; distinctive; disk-like. Fruit drupaceous; fleshy; not multiple; not splitting; enclosed in, or sitting on, the fleshy perianth; winged, or wingless. Seeds 1 per fruit; about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Stamens with two glands at the base; filaments opening by valves; ovary inferior; fruit in a cupule. Taxon images: hernand.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.