Emergent trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark white, cream, gray; not peeling in strips. Slash yellow, orange, red. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; encircling the twig; leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences represented by solitary flowers only; terminal (sometimes at the end of axillary short shoots). Flowers bisexual; stalked; about 0.5 cm long or across to to about 10 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 9-15. Stamens 100 (many); filaments present; free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; not awned; dehiscing by long slits. Ovary superior. Carpels separate, or partially joined, by base; 100 (many). Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit follicular; fleshy; multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Flowers sometimes from short shoots; gynoecium sessile. Taxon images: elmer.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.