Large trees to small trees. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots absent. Bark white, cream, gray, brown; not peeling in strips. Slash yellow, orange, brown. Vertical stripes in the inner bark or wood present, or absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense.
Leaves whorled; 3-5 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery, or paper-like; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules absent. Domatia present; pit-like; scattered along midrib. Basal glands present, or absent. Hairs on mature leaf present; dense.
Inflorescences present; terminal; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; to about 3 cm long or across, or to about 10 cm long or across; with one plane of symmetry; w