Small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; joined across twigs, or laterally placed; encircling the twig, or not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present, or absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals (4-)5; free, or partially joined; persistent in fruit; growing slightly after flowering. Epicalyx absent. Petals present; (4-)5; free; not fleshy; persistent; not stalked; entire. Stamens (8-)10; filaments present; free of each other; free of the perianth, or joined to the perianth (epi- and alternisepalous); all equal in size, or of different sizes; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; (2-)3. Locules (4-)6. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 4-20(-100) per fruit (many); less than 5 mm; winged; uniform in cross-section; aril absent; non-endospermic.
Leaves opposite with an intramarginal vein; sepals continue to grow after flowering; ovary inferior; ovary a half-inferior capsule. Taxon images: axinand.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.