Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent, aromatic. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves whorled; 4 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; on an unbranched axis; specialized inflorescence units present, or absent; specialized inflorescence units heads; numerous and persistent inflorescence bracts present at the bottom of the inflorescence. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals, or greatly reduced. Tepals 6 (unequal). Stamens 9; filaments present; free of each other; free of the perianth; all equal in size; number of stamens not a multiple of the adjacent perianth whorl. Anthers 4 loculed; not awned; dehiscing by valves. Ovary superior. Carpels solitary; 1. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; enclosed in, or sitting on, the fleshy perianth; wingless. Seeds 1 per fruit; less than 5 mm, or about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Leaves in whorls; perianth hairy; drupe on enlarged cup shaped perianth tube. Taxon images: genactin.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo – Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.