Large trees to shrubs. Buttresses present, or absent; short. Spines absent. Stilt roots present, or absent. Bark gray, brown; not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves spiral; not scale-like; compound; pinnate; without a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia present; pit-like; scattered along midrib, or scattered across lamina. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-5; free; not fleshy; deciduous; stalked, or not stalked; entire. Stamens (7-)8(-9); filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side, or attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2-3. Locules 2-3. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
1-2 leaflet pairs; fruits warty to spiny; seeds with sarcotesta. Taxon images: genxero.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.