Large trees to small trees. Buttresses present; short. Spines absent. Stilt roots absent. Bark white, green, gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves opposite; not scale-like; compound; pinnate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; winged, or not winged; attached to base of leaf blade; not swollen. Stipules present; joined; joined across twigs; encircling the twig; leafy; not fringed; large; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present; sparse.
Inflorescences present; axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present (reduced); the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4; free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4; free; not fleshy; deciduous; not stalked; entire. Stamens 8; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles free; 2. Stigmas 1 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2-100 per fruit (many); barely visible to less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Highland trees; leaves opposite, serrate, venation prominent; no glands on rachis; stipules interpetiolar; styles 2. Taxon images: weinmann.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.