Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple, or compound; pinnate; with a terminal leaflet; leaflets alternate; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end, or swollen at both ends. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 10; filaments present; joined, or free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles absent. Stigmas 2 lobed; distinctive; head shaped, or disk-like. Fruit capsular, or berry-like; fleshy, or non-fleshy; not multiple; splitting, or not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; about 1 cm long; wingless; uniform in cross-section; aril present (transparent and fleshy); non-endospermic.
Leaflet stalks swollen at rachis end; seeds surrounded by a fleshy aril. Taxon images: walsura.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.