Large trees to small trees. Buttresses present; short. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Slash cream, yellow, red, gray. Vertical stripes in the inner bark or wood absent. Odor resinous. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia present, or absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; persistent in fruit, or not persistent in fruit; not growing any further after flowering, or growing slightly after flowering, or with at least some sepals becoming wing-like; 5 becoming wing-like. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 15; filaments present; joined, or free of each other; free of the perianth; of different sizes; triplostemonous. Anthers 2 loculed; attached by the base; awned; dehiscing by long slits. Ovary superior, or partly inferior. Carpels joined; 3. Locules 3. Styles solitary. Stigmas 3 lobed; not distinctive. Fruit a nut; non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; winged, or wingless. Seeds 1 per fruit; about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Buttresses low and rounded; bark surface smooth, pale gray; calyx lobes valvate, about equal in length and generally shorter than the fruit. Taxon images: genvatic.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.