Large trees to small trees. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.
Leaves spiral; not scale-like; compound; bipinnate; without a terminal leaflet; leaflets opposite, or leaflets alternate; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present, or absent; 5; free; not fleshy; deciduous; stalked, or not stalked; entire. Stamens 8(-13); filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side to attached by the base; not awned; dehiscing by short slits. Ovary superior. Carpels joined; 3(-5). Locules 3(-5). Styles solitary. Stigmas 3 lobed; not distinctive. Fruit drupaceous; fleshy to non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-3 per fruit; less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.
Leaves bipinnate. Taxon images: tristiro.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.