Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous to leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined to wavy margined. Epidermis not waxy. Venation trinerved. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units present, or absent; specialized inflorescence units umbels, or corymbs; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4; free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4; free; fleshy; persistent; not stalked; entire. Stamens 4; filaments present; free of each other; free of the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 4. Locules 4. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 4 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves spiral, venation not very obvious; petioles slightly winged; flowers in parts of 4; filaments flattened; petals and sepals persistent; seeds oblong. Taxon images: tetramer.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.