Large trees to shrubs. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots present, or absent. Bark gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral (clustered in false whorls); not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery, or paper-like; gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible, or not visible. Smallest veins not prominent, but visible, or not visible; ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences represented by solitary flowers only; axillary. Flowers at least some flowers unisexual; with male and female flowers on different plants; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free, or partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free, or all joined (base); not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined (base); free of the perianth, or joined to the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2-3. Locules (1-)2-3(-6). Styles absent, or solitary. Stigmas 2-3 lobed; not distinctive. Fruit berry-like; fleshy, or non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2-100 per fruit (many); less than 5 mm to about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic (little), or non-endospermic.
Leaves spiral, but clustered at twig ends in pseudowhorls; anthers basifixed. Taxon images: ternstro.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.