Terminalia

Family Combretaceae. Order Myrtales.

Large trees to small trees. Buttresses present, or absent; tall. Spines present, or absent. Stilt roots absent. Bark yellow, brown; not peeling in strips. Slash yellow, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.

Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.

Leaves whorled, or spiral; 3-10 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; gland-dotted, or not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia present, or absent. Basal glands absent. Hairs on mature leaf absent.

Inflorescences present; terminal (rare), or axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on the same plant; not stalked, or stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals (4-)5; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens (8-)10; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary inferior. Carpels joined; 2-5 (reported for family). Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit drupaceous, or a samara; fleshy, or non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; winged, or wingless. Seeds 1 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.

Branches often in whorls at 90 degrees from trunk; leaves turn a striking yellow or red before falling. Taxon images: gentermi.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.