Large trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark red, gray, brown; not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences represented by solitary flowers only (rare), or present; terminal, or axillary; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; not stalked, or stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present, or absent. Perianth present; with distinct calyx and corolla whorls. Sepals 3-5; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3-5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 4-100 (many); filaments present; joined; joined to the perianth; all equal in size; isomerous, or diplostemonous, or triplostemonous, or polystemonous; opposite the sepals (when isomerous). Anthers 2 loculed; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 2-5. Locules 2-5. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; barely visible, or less than 5 mm; wingless; with folds like marble, in cross- section, or uniform in cross-section; aril absent; endospermic.
Leaves light green; margin toothed; calyx persistent; fruit bluish when ripe. Taxon images: gensymhe.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.