Small trees to shrubs. Buttresses absent. Spines present, or absent; on branches. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves two-ranked; not scale-like; simple; of different sorts on a single twig; not modified into spines; coriaceous, or leathery; gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined. Epidermis not waxy. Venation pinnate. Secondary veins open, or closed; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or axillary; the flowers from a single point on the stem, or on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts raised on a column; with the female parts of the flower raised on a stalk (short). Hypanthium absent. Disk present (enveloping ovary base); the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 3-5(-7); partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3-5(-7); free; not fleshy; deciduous; not stalked; entire. Stamens 5-15; filaments present; free of each other; free of the perianth; of different sizes (alternating long and short); isomerous, or diplostemonous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels solitary, or joined; 1-5. Locules 1-5. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-5 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves distichous and stiff; veins close and almost perpendicular to the midrib; often with stout thorns; disk enveloping ovary base. Taxon images: severina.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.