Large trees. Buttresses present; tall, or short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; disk-shaped, or star-like. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; dissected, or not dissected; palmately lobed; smooth margined. Epidermis not waxy. Venation pinnate, or palmate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein absent. Stalk of blade a quarter to one half the length of the blade to more than half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end (top). Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; large, or small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts raised on a column; with male and female parts raised on a stalk. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4-6; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 8; filaments present; joined; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; dehiscing by long slits. Ovary superior. Carpels joined; 2-5. Locules 2-5. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit follicular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; winged. Seeds 1-2 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Long petioles; palmate venation; flowers small; fruit with boat shaped wing; seed producing copious mucilage when ripe and wet. Taxon images: scaphium.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.