Small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts present at the bottom of the inflorescence. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 3; filaments present; free of each other; free of the perianth; all equal in size; anisomerous. Anthers 2 loculed; attached by the base; not awned; dehiscing by transverse slits. Ovary superior, or partly inferior. Carpels joined; 3. Locules 3. Styles absent. Stigmas 3 lobed; not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 18-100 per fruit (many); less than 5 mm; wingless; uniform in cross-section; aril present; endospermic.
Vegetative part with yellow 'pustules' - oil gland like; leaves alternate and entire; stipules barely visible; inflorescences with closely set decussate bracts; stamens 3; ovary 3-angled; stigma trilobed; 1-2 cm hairy bits appearing at the bottom of the seed. Taxon images: gensara.gif sarablum.gif sarawak.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.