Santiria

Family Burseraceae. Order Sapindales.

Large trees to small trees. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash cream, yellow, red. Vertical stripes in the inner bark or wood present, or absent. Odor absent. Exudate present, or absent; white, or colorless; spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.

Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.

Leaves spiral; not scale-like; compound; with three leaflets, or pinnate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.

Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 3; free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3; free; fleshy, or not fleshy; deciduous; not stalked; entire. Stamens 3, or 6; filaments present, or filaments absent; free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached along the side, or attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3. Locules 3. Styles absent, or solitary. Stigmas 1 lobed; distinctive, or not distinctive; disk-like. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril absent; non-endospermic.

Outer bark generally pale; inner bark laminated, sometimes hard mottled and granular; fruits brightly colored; stigma off center, nearly basal in fruit; seed round. Taxon images: gensanti.gif santiria.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.