Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present, or absent; T-shaped. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral, or two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent (and few). Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at both ends (very). Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary, or on the trunk or branches; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts present at the bottom of the inflorescence, or absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 3-4; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 5; filaments present; joined, or free of each other; joined to the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary superior. Carpels joined; 3-5 (3 or 5). Locules 1. Styles absent. Stigmas 2-5 lobed; distinctive, or not distinctive; disk-like. Fruit drupaceous; non-fleshy (leather); not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-6 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril absent; endospermic.
Slash smelling of cyanide; secondary veins few, spaced, prominent and strongly arching to the margin. Taxon images: genrypar.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.