Small trees to shrubs, or climbers. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs present, or absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple, or compound; pinnate; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at one end. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; free; persistent in fruit; growing slightly after flowering. Epicalyx absent. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 10; filaments present; joined; free of the perianth; of different sizes; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels separate; 4-5. Locules 1. Styles solitary. Stigmas 2 lobed; distinctive; head shaped. Fruit follicular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm, or about 1 cm long; wingless; uniform in cross-section; aril present; non-endospermic.
Apocarpous; stigma bilobed; either the testa fleshy or the seed covered by an arillode. Taxon images: genrour.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.