Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark brown; not peeling in strips. Exudate present; white; flowing; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves whorled; 3-10 per whorl; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open, or closed; not prominent, but visible. Smallest veins not prominent, but visible; net like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 5; filaments present; free of each other; joined to the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined, or partially joined, by styles; 2. Locules 2. Styles solitary. Stigmas 1 lobed; distinctive; head shaped. Fruit drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1-2 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves usually in whorls of three; carpels joined or not; fruit or fruitlets drupaceous with wart-like surface. Taxon images: rauv.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.