Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous to leathery; gland-dotted; symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin (rare). Epidermis not waxy. Venation pinnate. Secondary veins open; not prominent, but visible. Smallest veins not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; sparse.
Inflorescences present; axillary; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units present, or absent; specialized inflorescence units umbels; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on the same plant; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4-5; free, or partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 4-5; filaments absent; joined to the perianth; all equal in size; isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; not awned; dehiscing by long slits. Ovary partly inferior. Carpels joined; 4-5. Locules 1. Styles absent, or solitary. Stigmas 2-5 lobed; distinctive, or not distinctive; disk-like. Fruit berry-like; fleshy to non-fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; barely visible to less than 5 mm; wingless; with folds like marble, in cross- section (rare), or uniform in cross-section; aril absent; endospermic.
Filaments usually absent; style usually absent; fruit with one seed.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.