Large trees to small trees. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots present. Bark gray, brown; not peeling in strips. Slash white, cream, yellow. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves opposite; not scale-like; compound; pinnate, or bipinnate, or multiply compound; with a terminal leaflet; leaflets opposite; all of the same type; not modified into spines; coriaceous, or paper-like; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen, or swollen; swollen at both ends. Stipules absent. Domatia present, or absent; pit-like. Basal glands present, or absent. Hairs on mature leaf absent.
Inflorescences present; terminal, or on the trunk or branches; on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; with one plane of symmetry; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 2-5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 5; all joined; not fleshy; deciduous; not stalked; entire. Stamens 4-5; filaments present; free of each other; joined to the perianth; all equal in size; anisomerous, or isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles solitary. Stigmas 2 lobed; distinctive; club shaped. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm; winged; uniform in cross-section; aril absent; non-endospermic.
Leaves 1-3 times pinnate; glands often present at leaf bases; intramarginal vein present; calyx tube glandular/ warty; fruit with a corky central placenta. Taxon images: genrader.gif rader.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.