Large trees to small trees, or climbers. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash yellow, orange. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present; white; flowing; not blackening on exposure to air. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like, or box like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf absent.
Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units present; specialized inflorescence units heads; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; not stalked; with the parts barely visible to the eye; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 2-4 (?). Stamens 1; filaments present; anisomerous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels solitary. Locules 1. Styles solitary. Stigmas 2 lobed; not distinctive. Fruit fleshy; multiple; not splitting; enclosed in, or sitting on, the fleshy perianth; wingless. Seeds 100 per fruit (many); less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.
Flowers in heads and mixed in with numerous, variously shaped, bracts; style with two stigmas. Taxon images: prain.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.