Large trees, or small trees, or shrubs, or climbers. Buttresses absent. Spines absent. Stilt roots absent. Bark cream, gray, brown; not peeling in strips. Slash cream, gray, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base; not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate, or trinerved. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; box like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade, or a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; joined; laterally placed; encircling the twig; scale-like, or leafy; not fringed; large, or small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; terminal, or axillary; on a branched axis; specialized inflorescence units present; specialized inflorescence units heads; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; not stalked, or stalked; with the parts barely visible to the eye; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with all whorls looking like tepals. Tepals 2-4. Stamens 2-4; filaments present; free of each other; free of the perianth; all equal in size; isomerous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels solitary; 1. Locules 1. Styles absent, or solitary. Stigmas 1 lobed; distinctive, or not distinctive; head shaped, or disk-like. Fruit an achene; non-fleshy; multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); barely visible; wingless; uniform in cross-section; aril absent; endospermic.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.