Small trees to shrubs, or herbs. Buttresses absent. Spines present; on branches. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves two-ranked; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; very asymmetric at the base, or symmetric at the base (rare); not dissected; smooth margined. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent, or not prominent, but visible. Smallest veins not prominent, but visible, or not visible; net like. Intramarginal vein present, or absent. Stalk of blade absent to less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences present; axillary, or on the trunk or branches; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant; stalked; with the parts barely visible to the eye to about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present (3-6 glands); the stamens inside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 6; free, or partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 3-6; filaments present; joined, or free of each other; free of the perianth; all equal in size, or of different sizes; anisomerous, or isomerous; alternating with the sepals (when isomerous). Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 3(-8). Locules 3. Styles solitary (rare), or free; 3. Stigmas 1-3 lobed (1 or 3); not distinctive. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 6 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves asymmetric and distichous; flowers tiny and appearing in leaf axils. Taxon images: phyllan.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.