Small trees to shrubs. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or wavy margined, or with small to large teeth on margin. Epidermis waxy, or not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; ladder like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia present, or absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; axillary; on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; with the parts barely visible to the eye, or about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens on the disk. Perianth present; with distinct calyx and corolla whorls. Sepals 4-8; free, or partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 4-5; free; not fleshy; deciduous; not stalked; entire. Stamens 4-5(-8); filaments present; free of each other; free of the perianth; all equal in size; isomerous, or diplostemonous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 2. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 2-4 per fruit; less than 5 mm; wingless; uniform in cross-section; aril present; endospermic.
Shrubs or small trees; branchlets zig-zag; leaves distichous; midrib prominent; domatia sometimes present. Taxon images: genperro.gif perrott.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.