Emergent trees to shrubs. Buttresses present, or absent; tall. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; star-like. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves opposite; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present; free from one another; joined across twigs; encircling the twig; leafy; not fringed; small; persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present, or represented by solitary flowers only; axillary, or leaf-opposed; the flowers from a single point on the stem, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium present. Disk present, or absent. Perianth present; with distinct calyx and corolla whorls. Sepals 3-6; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 3-6; free; not fleshy; deciduous; stalked; lobed, or fringed like a comb. Stamens 6, or 8, or 10, or 12; filaments present; joined, or free of each other; joined to the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary inferior. Carpels joined; 9-12. Locules 9-12. Styles solitary. Stigmas 8-10 lobed; distinctive; head shaped. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); barely visible, or less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.
Leaves opposite; twigs hollow; sepals joined. Taxon images: genpella.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.