Payena

Family Sapotaceae. Order Ebenales.

Large trees to small trees. Buttresses absent. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash yellow, red. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate present; white; flowing, or spotty; not blackening on exposure to air. Terminal buds not enclosed by leaves.

Complex hairs present, or absent; disk-shaped. Stinging hairs absent. Mature twig indumentum absent.

Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open, or closed; prominent. Smallest veins prominent, or not prominent, but visible; net like. Intramarginal vein present. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules present, or absent; free from one another; laterally placed; not encircling the twig; scale-like; not fringed; small; not persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.

Inflorescences present, or represented by solitary flowers only; axillary, or leaf-opposed; the flowers from a single point on the stem; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 4; free; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; 7-10; all joined; not fleshy; deciduous; not stalked; entire. Stamens 14, or 16, or 20(-100) (many); filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side, or attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; non-endospermic.

Leaves spaced along twigs; petioles of even thickness and grooved; intramarginal vein present. Taxon images: genpayen.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.