Large trees to small trees. Buttresses present, or absent; tall, or short. Spines absent. Stilt roots absent. Bark red, gray, brown; not peeling in strips. Slash brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; dense.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation palmate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade more than half the length of the blade; not winged; attached to surface of leaf blade to attached to base of leaf blade; swollen; swollen at one end. Stipules absent. Domatia absent. Basal glands absent. Hairs on mature leaf present, or absent; dense, or sparse.
Inflorescences represented by solitary flowers only (female), or present (male); axillary; on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on the same plant, or with male and female flowers on different plants; stalked; to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present. Perianth present; with distinct calyx and corolla whorls. Sepals 0; fully joined; joined together and forming a lid; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; (4-)5-8(-9); free; not fleshy (but each with a large hairy scale-like appendage); deciduous; not stalked; entire. Stamens 20-25(-100) (many); filaments present; free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2-4. Locules 1. Styles absent. Stigmas 2-4 lobed; not distinctive. Fruit berry-like; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); much more than 1 cm long; wingless; uniform in cross-section; aril present (white and fleshy); endospermic.
Large trees; leaves with palmate venation, bases heart shaped; petioles long; fruits large, with large kidney shaped seeds. Taxon images: pangium.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.