Emergent trees to large trees. Buttresses present; tall. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Slash cream, yellow, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present, or absent; disk-shaped. Stinging hairs absent. Mature twig indumentum absent.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; gland-dotted, or not gland-dotted; symmetric at the base; not dissected; smooth margined, or with small to large teeth on margin. Epidermis not waxy. Venation palmate. Secondary veins open; prominent. Smallest veins not prominent, but visible; ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia present; pit-like; scattered along midrib. Basal glands absent. Hairs on mature leaf present; sparse.
Inflorescences present; on the trunk or branches (branches); on an unbranched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers at least some flowers unisexual; with male and female flowers on different plants; not stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5-8; free; not persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals absent. Stamens 5-8; filaments present; free of each other; free of the perianth; all equal in size; isomerous; opposite the sepals (when isomerous). Anthers 2 loculed; attached along the side, or attached by the base; not awned; dehiscing by short slits, or dehiscing by long slits. Ovary inferior. Carpels joined; 5-8. Locules 1. Styles free; 5-8. Stigmas 1 lobed; distinctive; head shaped. Fruit capsular; non-fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); barely visible; wingless; uniform in cross-section; aril absent; non-endospermic.
Large, very fast growing trees; bark light gray and prominently hooped; leaves large and rounded.
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.