Nyssa

Family Nyssaceae. Order Cornales.

Emergent trees to large trees. Buttresses present, or absent; short. Spines absent. Stilt roots absent. Bark gray; not peeling in strips. Slash yellow, brown. Vertical stripes in the inner bark or wood present, or absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.

Complex hairs absent. Stinging hairs absent. Mature twig indumentum present, or absent; sparse.

Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent; net like. Intramarginal vein present, or absent. Stalk of blade less than a quarter the length of the blade; not winged; attached to base of leaf blade; not swollen. Stipules absent. Domatia present, or absent. Basal glands absent. Hairs on mature leaf present; sparse.

Inflorescences present; axillary; on an unbranched axis; specialized inflorescence units present; specialized inflorescence units heads; numerous and persistent inflorescence bracts absent. Flowers bisexual, or at least some flowers unisexual; with male and female flowers on the same plant; not stalked; about 0.5 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk present; the stamens outside the disk. Perianth present; with distinct calyx and corolla whorls. Sepals (4-)5; partially joined; persistent in fruit; not growing any further after flowering. Epicalyx absent. Petals present; (4-)5-8; free; not fleshy; deciduous; not stalked; entire. Stamens 8-10; filaments present; free of each other; free of the perianth; all equal in size; diplostemonous. Anthers 2 loculed; attached along the side; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 2. Locules 1. Styles solitary. Stigmas 1 lobed; not distinctive. Fruit berry-like, or drupaceous; fleshy; not multiple; not splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 1 per fruit; less than 5 mm; wingless; uniform in cross-section; aril absent; endospermic.

Trunk and twig bark with prominent lenticels; leaves bunched at twig ends; flowers in stalked heads, male and hermaphrodite; style spliiting into two stigmatic branches. Taxon images: nyssa.gif


Cite this publication as: Jarvie, J.K. and Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and Illustrations' URL http://django.harvard.edu/users/jjarvie/Borneo.htm/.
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.