Large trees to small trees. Buttresses present; short. Spines absent. Stilt roots absent. Bark gray, brown; not peeling in strips. Slash red, brown. Vertical stripes in the inner bark or wood absent. Odor absent. Exudate absent. Terminal buds not enclosed by leaves.
Complex hairs present; disk-shaped, or star-like. Stinging hairs absent. Mature twig indumentum present, or absent; dense, or sparse.
Leaves spiral; not scale-like; simple; all of the same type; not modified into spines; coriaceous, or leathery; not gland-dotted; symmetric at the base; not dissected; smooth margined. Epidermis not waxy. Venation pinnate. Secondary veins open; prominent. Smallest veins prominent, or not prominent, but visible; net like, or ladder like. Intramarginal vein absent. Stalk of blade less than a quarter the length of the blade to a quarter to one half the length of the blade; not winged; attached to base of leaf blade; swollen; swollen at one end. Stipules present; free from one another; laterally placed; not encircling the twig; scale-like, or leafy; not fringed; large; not persistent, or persistent. Domatia absent. Basal glands absent. Hairs on mature leaf present; dense, or sparse.
Inflorescences present; axillary; the flowers from a single point on the stem, or on an unbranched axis, or on a branched axis; specialized inflorescence units absent; numerous and persistent inflorescence bracts absent. Flowers bisexual; stalked; about 0.5 cm long or across, or to about 3 cm long or across; round; with male or female parts not raised on a column. Hypanthium absent. Disk absent. Perianth present; with distinct calyx and corolla whorls. Sepals 5; partially joined; not persistent in fruit; not growing any further after flowering. Epicalyx present. Petals present; 5; free; not fleshy; deciduous; not stalked; entire. Stamens 100 (many); filaments present; joined, or free of each other; free of the perianth; all equal in size; polystemonous. Anthers 2 loculed; attached by the base; not awned; dehiscing by long slits. Ovary superior. Carpels joined; 5. Locules 5. Styles solitary. Stigmas 5 lobed; distinctive; head shaped. Fruit capsular; fleshy; not multiple; splitting; not enclosed in, or sitting on, fleshy perianth parts; wingless. Seeds 100 per fruit (many); less than 5 mm, or about 1 cm long, or much more than 1 cm long; wingless; uniform in cross-section; aril present; non-endospermic.
Parts covered in stellate hairs of fringed scales; petiole long and kneed; leaves generally obovate, notched or rounded above; stipules large; flower bud enclosed by epicalyx; fruit tesselate, splitting before falling, inside covered in irritant hairs. Taxon images: genneesi.gif
Cite this publication as: Jarvie, J.K. and
Ermayanti (1996 onwards). `Tree Genera of Borneo - Descriptions and
Dallwitz (1980) and Dallwitz, Paine and Zurcher (1993) should also be cited.